Professional Titles

• Assistant Director, Biological Sciences
• Curator Emeritus, Department of Recent Invertebrates
• Research Associate, American Museum of Natural History
• Research Associate, Florida State Collection of Arthropods
• Research Associate, Museu d'histoire Naturelle, Geneva, Switzerland

Education

• Ph.D. in Zoology, Virginia Polytechnic Institute & State University, 1960
• M.A. in Entomology, 1956, Cornell University
• University of Virginia, 1945-1950

 Professional Experience

• Curator of Recent Invertebrates, Virginia Museum of Natural History
• Professor of Biology, Radford University
• NSF Research Associate, United States National Museum of Natural History
• Graduate Research Assistant, Virginia Polytechnic Institute & State University
• Chemist, Radford Army Ammunition Plant, Radford
• Research Technician, Virginia Fisheries Laboratory, Yorktown

Research Interests

• Biota of the Southern Appalachians
• Natural history of Virginia
• Systematics and global biogeography of Diplopoda, Chilopoda, and Arachnida (especially East African and Brasilian millipeds)
• Distribution and biology of Virginia arthropods (especially myriapods, larger crustacea, arachnids, heteroptera,
  carabid, longhorned beetles, aquatic insect orders)
• Distribution and systematics of Virginia freshwater mollusks (especially unionids and pleurocerids)

Recent Professional Service

• Curator of Recent Invertebrates, Virginia Museum of Natural History
• Professor of Biology, Radford University
• NSF Research Associate, United States National Museum of Natural History
• Graduate Research Assistant, Virginia Polytechnic Institute & State University
• Chemist, Radford Army Ammunition Plant, Radford
• Research Technician, Virginia Fisheries Laboratory, Yorktown

Selected Publications

• 1958. "Checklist of the Millipeds of North America," with R. V. Chamberlin, United States National Museum Bulletin. 212, pp. 1-236.
• 1963. "A revision of the North American annelid worms of the genus Cambarincola (Oligochaeta:Branchiobdellidae)." Proceedings of the. United States National Museum 114:271-371.
• 1969. "Myriapoda, exclusive of Insecta," in: Treatise of Invertebrate Palaeontology,Vol. R, pp. R573-606.
• 1980. Classification of the Diplopoda. Museum d'Histoire Naturelle, Geneva, Switzerland. 237 pp.
• 1982. "Chilopoda" and "Diplopoda" sections (Vol. 2, pp. 681-724) in: S. B. Parker, ed., Synopsis and Classification of Living  Organisms. McGraw-Hill, New York.
• 1990. "Diplopoda," pp. 835-860, in : D. L. Dindall, et., Soil Biology Guide. Wiley Interscience, New York. 1349 pp.
• 1990 "Myriapoda 5. Polydesmida 4. Family Oxydesmidae," in: Das Tierreich, W. DeGruyter & Co., Berlin & New York, part 107. pp. 1-512.
• 1991. Chapters "The Nature of Virginia" (pp. 23-48) and "Arthropoda" (pp. 173-249), in: Virginia's Endangered Species, McDonald & Woodward Publishing Co., Blacksburg, VA
• 1996 Seed Bugs of Virginia. The Insects of Virginia. Part 14. Virginia Museum of Natural History, Martinsville. 111 pp.
• 1999 Checklist of the Millipeds of North and Middle America. Special Publication 8, Virginia Museum of Natural History, Martinsville. 584 pp.
• 2002. The Gomphodesmidae: An African Family of Polydesmoid Diplopoda. Naturhistorisches Museum, Vienna, Austria (in press).
• 2002. Assassin Bugs of Virginia. The Insects of Virginia, Part 15. Virginia Museum of Natural History, Martinsville (in press).